When the "compatibility with common mapping errors" options were introduced, the idea was that they should not apply when recording, during playback or whenever using a "vanilla" complevel (<7).
However, I notice that comperr_zerotag (at least) applies during recording and playback and with vanilla complevels.
Example demo and wad:
In this demo I deliberately exploit a missing tag error. If comperr_zerotag = 1, the demo desyncs.
I haven't tested the other comperr setting to see if they suffer from the same problem.
The earliest version I have with the bug is 2012/01/28, and the latest version without it is 2011/12/04. I presume that something in r4156 is to blame.