Re: [Nemo2-user] Nemo2: question on population stat outputs
Individual-based forward-time genetics simulation software
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From: Francois L. (PACA) <fra...@in...> - 2020-07-08 15:45:35
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Dear Fred, many thanks for your quick answer, much appreciated too! Yes it helps! For 1/ this is what I understood For 2/ OK I will disregard those migration stats. For final clarification on this 2nd point: as you correctly understood there is both pollen and seed dispersal (different migration rates), do you mean that in such case the migration stats cumulate pollen grains and zygotes (seeds) together? Best, François Le 08/07/2020 à 16:06, Fred Guillaume a écrit : > Dear François, > > thanks for posting to the list, this is very much appreciated! > > the answers to your questions are: > > 1/ the *regulation* LCE independently regulates the offspring and adults > to the patch capacities - this is indeed why you have K offspring AND K > adults after regulation. > Note that regulation is also performed in the *aging* LCE where max K > offspring are chosen to become the new adult generation in each patch. > You may thus not need to add *regulation* in your life cycle. > > 2/ the number of residents (and other migration stats) is calculated > during migration, i.e., when the *disperse* LCE (or *breed_disperse* as > in your case) is executed in the life cycle. Therefore, the number of > residents depends on the number of offspring when dispersal is processed > and not when you record the stats (with *save_stats*). resid.pi will > exceed the number of adults when fecundity is high and migration rates > are low. > > Now, your case is special because you mix forward (seed) and backward > (pollen) migration. Thus, residents are counted during each event and > will largely exceed the carrying capacities. In your case I would > disregard those stats, or you can place *save_stats* inbetween the two > dispersal LCEs to isolate their effect on the migration counters. Note > that for backward migration, a resident is a pollen grain that > originated from a local parent. In that case, each parent (adult) can be > both a resident or a migrant depending on where it sends its pollen grains. > > I hope this helps! > > Best, > Fred > > > On 7/8/20 2:56 PM, Francois Lefevre (PACA) wrote: >> Hello, >> (sorry if this mailing list is not the appropriate way to post these >> questions, if it is not just tell me) >> >> two questions about the population stat outputs: >> >> 1) for each patch /i/, both statistics of the nb of offspring >> (/off.fem.pi)/ and nb of adults (/adlt.fem.pi)/ can reach the patch >> capacity eventhough they are computed after the regulation step (see >> Life cycle below): do I correctly understand that population regulation >> is made independently for offspring and adults? >> (after aging and mortality of the adults, only the younger cohort is >> kept and becomes adult, ending to the appropriate capacity) >> >> 2) how is computed the number of residents per patch (/resid.pi)/ ? This >> number largely exceeds the carrying capacity and also exceeds the number >> of adults of the patch: does it represent the "memory" of the total >> number of local individuals before regulation, i.e. nb of adults and >> locally produced offspring? >> >> The Life cycle is the following: >> breed_disperse 1 >> seed_disp 2 >> viability_selection 3 >> regulation 4 >> save_stats 5 >> store 6 >> aging 7 >> save_files 8 >> >> Thank you! >> François >> >> -- >> François Lefèvre >> INRAE, Ecologie des Forêts Méditerranéennes, URFM, 84914 Avignon, France >> Domaine Saint Paul, 228 route de l'Aérodrome Agroparc >> +33 (0)4 32 72 29 01 >> >> >> >> _______________________________________________ >> Nemo2-user mailing list >> Nem...@li... >> https://lists.sourceforge.net/lists/listinfo/nemo2-user >> -- François Lefèvre INRAE, Ecologie des Forêts Méditerranéennes, URFM, 84914 Avignon, France Domaine Saint Paul, 228 route de l'Aérodrome Agroparc +33 (0)4 32 72 29 01 |