Re: [Nemo2-user] Nemo2: question on population stat outputs

[Fred G. <gui...@gm...>]

Dear François,

thanks for posting to the list, this is very much appreciated!

the answers to your questions are:

1/ the *regulation* LCE independently regulates the offspring and adults
to the patch capacities - this is indeed why you have K offspring AND K
adults after regulation.
Note that regulation is also performed in the *aging* LCE where max K
offspring are chosen to become the new adult generation in each patch.
You may thus not need to add *regulation* in your life cycle.

2/ the number of residents (and other migration stats) is calculated
during migration, i.e., when the *disperse* LCE (or *breed_disperse* as
in your case) is executed in the life cycle. Therefore, the number of
residents depends on the number of offspring when dispersal is processed
and not when you record the stats (with *save_stats*). resid.pi will
exceed the number of adults when fecundity is high and migration rates
are low.

Now, your case is special because you mix forward (seed) and backward
(pollen) migration. Thus, residents are counted during each event and
will largely exceed the carrying capacities. In your case I would
disregard those stats, or you can place *save_stats* inbetween the two
dispersal LCEs to isolate their effect on the migration counters. Note
that for backward migration, a resident is a pollen grain that
originated from a local parent. In that case, each parent (adult) can be
both a resident or a migrant depending on where it sends its pollen grains.

I hope this helps!

Best,
Fred


On 7/8/20 2:56 PM, Francois Lefevre (PACA) wrote:
> Hello,
> (sorry if this mailing list is not the appropriate way to post these
> questions, if it is not just tell me)
> 
> two questions about the population stat outputs:
> 
> 1) for each patch /i/, both statistics of the nb of offspring
> (/off.fem.pi)/ and nb of adults (/adlt.fem.pi)/ can reach the patch
> capacity eventhough they are computed after the regulation step (see
> Life cycle below): do I correctly understand that population regulation
> is made independently for offspring and adults?
> (after aging and mortality of the adults, only the younger cohort is
> kept and becomes adult, ending to the appropriate capacity)
> 
> 2) how is computed the number of residents per patch (/resid.pi)/ ? This
> number largely exceeds the carrying capacity and also exceeds the number
> of adults of the patch: does it represent the "memory" of the total
> number of local individuals before regulation, i.e. nb of adults and
> locally produced offspring?
> 
> The Life cycle is the following:
> breed_disperse 1
> seed_disp 2
> viability_selection 3
> regulation 4
> save_stats 5
> store 6
> aging 7
> save_files 8
> 
> Thank you!
> François
> 
> -- 
> François Lefèvre
> INRAE, Ecologie des Forêts Méditerranéennes, URFM, 84914 Avignon, France
> Domaine Saint Paul, 228 route de l'Aérodrome Agroparc
> +33 (0)4 32 72 29 01
> 
> 
> 
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