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Metamodels provide a novel approach to species risk assessment in which diverse threats are addressed through different disciplines, act at different spatiotemporal scales, or interact in non-linear ways. A metamodel links multiple, discrete models that depict components of a complex system, governing the flow of information between models and the sequence of events in the simulation. Each discrete model simulates processes specific to its disciplinary realm while being informed of changes in other parts of the metamodel by accessing common descriptors of the system, populations, and individuals. ...